It is the short form of:
When in Rome do as the Romans do
In Japanese, you'd say 郷に入っては、郷に従え (ごうにいってはごうにしたがえ). We use this to say that when you are in a new environment you should do what the local people are doing. It is such a common saying that it is often shortened to "When in Rome..."
The most common usage is you are travelling or somewhere new with your friend and suddenly you see some strange or different behavior and are not sure if you want to join in and try something new.
Al: What kind of meat is this?
Bill: I think it's whale meat...
Al: Whale!? I don't want to eat whale!
Bill: When in Rome...
Al: Well, ok. I'll try a bite.
----
Craig: What!?! Some kid just sprayed water on me!
Sam: Oh, I think it's Songkran the water festival. Just spray him back!
Craig: Well, you know what they say: "When in Rome..."
Sam: "...do as the Romans do!"
Have a nice Golden Week,
Phil
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